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Invalid Recipient 5.1.1

Hello, I can send via outlook or cpanel webmail (Round Cube) with an email i recently setup....i can send without issues..when replying it gives a bounceback from the correct server (that hosts my domain and mail) but says recipient is invalid, please check, etc.

 

Any ideas why?  DNS records are pointing internally, cpanel email mx entry is local, not remote.

For some reason its not seeing my account on inbound as legitimate?

 

Thank You.